Hercules-club (also known as Prickly Ash, Toothache Tree, or Devil's Walking Stick)
Kingdom: Plantae, Division: Tracheophyta, Class: Magnoliopsida, Order: Sapindales, Family: Rutaceae, Genus: Zanthoxylum, Species: Z. clava-herculis · Rutaceae (Citrus or Rue family) · Pinnately compound, deciduous

Species
Zanthoxylum clava-herculis
Leaf Type
Pinnately compound, deciduous
Family
Rutaceae (Citrus or Rue family)
Shape
Elliptical to ovate leaflets, arranged in pairs along a rachis with a terminal leaflet
Size
Individual leaflets are approximately 1.5 to 3 inches long; the overall compound leaf can reach 8 to 15 inches in length, typical for mature specimens
Margin
Crenate to serrate (finely toothed margins)
Arrangement
Alternate arrangement on the woody stem
Rarity
Common within its native southeastern U.S. range, though less frequently encountered in northern pockets of its distribution
Color & Pattern
Medium to dark green upper surface with lighter green undersides; the leaflets in this image display minor chlorotic yellow spotting between veins, likely due to nutritional stress or insect feeding
Venation Pattern
Pinnate venation with a prominent central midrib and branching lateral veins on each leaflet
Texture & Surface
Smooth (glabrous) surface with a slightly leathery texture; the rachis often features small prickles
Description
Hercules-club is a small to medium-sized deciduous tree or large shrub known for its exceptionally stout, sharp prickles that stud its grey bark and stems. It produces large, compound leaves with glossy green leaflets, clusters of small greenish-white flowers in spring, and small, dark fruits that attract wildlife
Key Features
Extremely sharp, conical prickles mounted on corky bases covering the trunk and branches, combined with large, pinnately compound leaves featuring finely toothed, aromatic leaflets
Habitat
Found primarily in moist soils of bottomland forests, river floodplains, swamp borders, and occasionally dry upland woods; prefers full sun to partial shade
Geographic Range
Native to the southeastern and south-central United States, ranging from Texas and Oklahoma east to Florida, and north to Virginia and Illinois
Condition Notes
Fair condition; the leaflets show visible chlorotic mottling (yellowish spots) and minor insect herbivory along the edges, but the overall structural integrity of the compound leaf remains intact
Interesting Facts
The common name 'Toothache Tree' comes from the traditional use of its bark and leaves to numb mouth pain due to a numbing chemical compound found in the plant, which causes a tingling sensation when chewed
Medicinal & Uses
Historically used by Native Americans and early settlers as a remedy for toothaches, rheumatism, and digestive issues due to its local anesthetic and stimulant properties; sometimes planted in wildlife gardens
Ecological Role
Serves as an important host plant for the larvae of the Giant Swallowtail butterfly (Papilio cresphontes) and provides berries that are consumed by numerous bird species
Similar Species
Can be confused with other prickly plants or sumacs (Rhus species), but can be distinguished by its distinct citrus-family aroma when crushed and the unique woody prickles directly on the bark and leaf rachis
Foraging Tips
Look in moist bottomlands or forest edges within the American South; always check for the characteristic sharp prickles on the trunk before handling, and crush a leaf gently to test for the signature citrus-like or pungent scent
Toxicity Warning
Non-toxic, though the plant contains active alkaloids that cause a strong numbing sensation in the mouth if chewed, and the sharp prickles can cause physical punctures and irritation